新疆准噶尔盆地玛湖凹陷玛页...拉尔统风城组孢粉地层学研究_师天明.pdf

收稿日期:2021-12-21;改回日期:2022-06-21;录用日期:2022-06-26*中国科学院战略性先导科技专项(B 类)(XDB26010501)、中国科学院前沿科学重点研究项目(QYZDY-SSW-DQC023)、第二次青藏高原综合科学考察研究(2019QZKK0706)、国家自然科学基金委地区基金重点项目(U1702242)和新疆油田公司项目(2020-C4011)联合资助。*通讯作者:彭辉平,工程师,e-mail: 古生物学报,61(4):615627(2022年12月)Acta Palaeontologica Sinica,61(4):615627(December 2022)研究论文 DOI:10.19800/ki.aps.2021075 新疆准噶尔盆地玛湖凹陷玛页1井宾夕法尼亚亚系 至乌拉尔统风城组孢粉地层学研究*师天明1 张 华2 肖继南1 阿木提阿丽亚1 刘 锋2 唐 鹏2 朱莉叶2 谢 媛2 彭辉平2*1 中国石油新疆油田分公司实验检测研究院,新疆克拉玛依 834000;2 中国科学院南京地质古生物研究所,现代古生物学和地层学国家重点实验室,南京 210008 提要 准噶尔盆地西北缘玛湖凹陷风城组主要形成于碱湖环境,含有优质生烃母岩。对玛页1井风城组岩心样品的孢粉分析建立了Protohaploxypinus perfectusLunatisporites tersus(PT)孢粉组合。该组合包括20属29种孢粉化石。PT组合以双气囊具肋花粉占主导,蕨类孢子含量很低为特征;孢粉母体植物类群以裸子植物门种子蕨盾籽目 为 主,其 次 为 松 柏 纲 松 柏 目。该 组 合 与 准 噶 尔 盆 地 南 缘 塔 什 库 拉 组 上 部 至 乌 拉 泊 组 的CrustaesporitesProtohaploxypinusHamiapollenites孢粉组合可以对比,均以双气囊具肋花粉为主要特征,又同时出现重要的属种Gardenasporites bilabiatus,Triangulisaccites boleensis和Hamiapollenites saccatus。孢粉地层学和同位素年代学资料表明,玛页1井风城组PT组合的时代很可能属于石炭纪宾夕法尼亚亚纪卡西莫夫期至二叠纪乌拉尔世阿瑟尔期,玛湖凹陷区整个风城组沉积时代晚于宾夕法尼亚亚纪巴什基尔期,其上部可能包含部分乌拉尔世阿瑟尔期沉积。风城组黑色页岩中产出的双气囊具肋花粉占绝对优势的孢粉组合和冷水古鳕类化石,与互层状盐碱层中自生矿物碳镁钠石和碳酸钠钙石共同表明风城组页岩和盐碱韵律沉积很可能形成于冷干和暖干频繁交替的古环境中。关键词 风城组 孢粉学 宾夕法尼亚亚系 乌拉尔统 准噶尔盆地 古气候 中文引用 师天明,张华,肖继南,阿木提阿丽亚,刘锋,唐鹏,朱莉叶,谢媛,彭辉平,2022.新疆准噶尔盆地玛湖凹陷玛页1井风城组孢粉地层学研究.古生物学报,61(4):615627.DOI:10.19800/ki.aps.2021075 英文引用 Shi Tian-ming,Zhang Hua,Xiao Ji-nan,Amuti Aliya,Liu Feng,Tang Peng,Zhu Li-ye,Xie Yuan,Peng Hui-ping,2022.Palynostratigraphy of the Fengcheng Formation(Pennsylvanian to Cisuralian)from the Maye-1 bore-hole in the Mahu sag,northwestern Junggar Basin,Xinjiang,Northwest China.Acta Palaeontologica Sinica,61(4):615627.DOI:10.19800/ki.aps.2021075 Palynostratigraphy of the Fengcheng Formation(Pennsylvanian to Cisuralian)from the Maye-1 borehole in the Mahu sag,northwestern Junggar Basin,Xinjiang,Northwest China SHI Tian-ming1,ZHANG Hua2,XIAO Ji-nan1,AMUTI Aliya1,LIU Feng2,TANG Peng2,ZHU Li-ye2,XIE Yuan2,PENG Hui-ping2 616 古 生 物 学 报 61 卷 1 Research Institute of Experiment and Detection of Xinjiang Oilfield Company,Karamay,834000,Xinjiang,China;2 State Key Laboratory of Palaeobiology and Stratigraphy,Nanjing Institute of Geology and Palaeontology,Nanjing 210008,China Abstract The Fengcheng Formation in the Mahu sag in northwestern Junggar Basin,Xinjiang,was deposited in an alkaline saline lake and contains large volumes of high-quality source rocks.PetroChina Xinjiang Oilfield Company conducted the Maye-1 borehole project and collected the entire sequence of the Fengcheng Formation in the Mahu sag to evaluate and explore the oil and gas potential in this area.In order to better constrain the age of the Fengcheng For-mation,a palynostratigraphic project has been conducted since 2019.A total of fifty palynological samples,mainly of shale,claystone and siltstone,were collected from the Maye-1 core.All samples were cleaned,crushed,weighed(30-50 g for each sample),and macerated following the standard HCl-HF-HCl method.The organic residues were sieved through 180 and 10 m meshes,respectively.The residues were then mounted on microscope slides using glyc-erin jelly.Palynomorphs were observed and photographed using a LEICA DM 2500 microscope equipped with a D800E camera.However,a considerable number of palynomorphs are not well preserved,usually with various sizes of secondary cavities or perforations.All slides are stored in the PetroChina Xinjiang Oilfield Company,Karamay,Xin-jiang,China.Positions of the spores and pollen illustrated in the text figures are located using an England Finder Slide.Based on systematic palynological identification and statistics,a new fossil palynological assemblage,Protohaploxy-pinus perfectus-Lunatisporites tersus(PT)assemblage,is established for the Fengcheng Formation.Twenty-nine spe-cies in twenty genera of spores and pollen were recognized in the PT Assemblage from fifty macerated samples.In the PT assemblage,the Disaccites striatiti pollen is dominant,while the fern spores are relatively rare.The comparison of the dispersed and in-situ spores and pollen associated with their reliable parent plants shows that the palynoflora con-sists of dominant species of Peltaspermales and species of Coniferales as the second major component.The PT assem-blage is correlatable with the CrustaesporitesProtohaploxypinusHamiapollenites assemblage from the upper part of the Tashenkula Formation to the Wulabo Formation at the southern margin of the Junggar basin due to the high portion of the Disaccites striatiti pollen in the palynoflora and the crucial co-occurring species,including Gardenasporites bi-labiatus,Triangulisaccites boleensis and Hamiapollenites saccatus.In addition,the PT Assemblage,mainly consisting of the Disaccites striatiti pollen(70%),differs from the Bashkirian Calamospora breviradiataLunatisporites tersus assemblage from the age-constrained Jiamuhe Formation in the nearby Zhongguai uplift.The latter assemblage bears a slightly less Disaccites striatiti pollen(ca.60%)and an equal amount of Calamospora spores(ca.40%),indicating that the PT assemblage is reliably dated as post-Bashkirian.Based on the precise U-Pb zircon dates from two tuff layers(304.9 0.69 Ma and 299.9 0.64 Ma)in the lower part and a tuff bed(296.9 0.8 Ma)in the upper part of the Fengcheng Formation in the Mahu sag,the age of Fengcheng Formation is constrained to the KasimovianAsselian.In summary,careful palynological comparisons and precise geochronological data suggest that the PT assemblage from the Fengcheng Formation of the Maye-1 borehole was likely deposited from the Kasimovian to the Asselian age.Moreover,the age of the entire Fengcheng Formation in the Mahu sag should be younger than the Bashkirian,and the upper part of the Fengcheng Formation may contain some Asselin sediments.Abundant Disaccites striatiti pollen and the intriguing emergence of the cold-water Gadus in the shales and the warm-phase authigenic alkali minerals of eitelite and shortite from the interbedded saline layers indicate that the rhythmic sedimentary strata of the Fengcheng Formation in the Mahu sag were probably formed in paleoenvironments with frequent fluctuations of cold-dry and warm-dry paleoclimate conditions.Keywords Fengcheng Formation,palynology,Pennsylvanian,Cisuralian,Junggar Basin,paleoclimate 1 前 言 新疆准噶尔盆地是中国重要的含油气盆地(王小军等,2021;张元元等,2021;Cao et al.,2020等)。为了进一步提高该盆地石油产出量,中国石油集团新疆油田公司不断拓展盆地的油气勘探开发区块。为了准确寻找具有开发价值的目标油气储集层和生油气层,该公司需要对研究新区的地层开展研究,建立地层层序框架,了解地层的分布,为新区的油气勘探提供地层基础。玛页1井对风城组的钻探为新疆准噶尔盆地西北缘玛湖凹陷地区的油气资源勘探拉开了序幕(Wang et al.,2021)。迄今,对风城组年代的认识还存在较大争议。路玉(2018)从风城组碎屑锆石中获得了绝对年龄277.4278.9 Ma的结果,认为风城组沉积于二叠纪乌拉尔世。这一观点被普遍采纳(例 4期 师天明等:新疆玛湖凹陷孢粉地层学;Shi et al.:Palynostratigraphy of Mahu Sag,Xinjiang 617 如:魏研等,2021;Yu et al.,2018,2019;Gao et al.,2018等)。但是,也有一些学者认为风城组的时代跨越了石炭二叠纪界线,其底部可能存在石炭纪宾夕法尼亚亚纪的地层。如:Cao等(2020)认为风城组沉积于冈瓦纳大陆晚古生代第三次大冰期期间(约305297 Ma);Wang等(2021)在风城组火山灰碎屑锆石中获得绝对年龄范围为304.9296.6 Ma,指示该组绝大部分沉积于宾夕法尼亚亚纪。另外,关于风城组形成的古环境背景也存在一定分歧。余宽宏等(2016)使用18O/13C比值判断气候的方法,认为风城组含碱韵律层形成于温暖潮湿和干旱炎热频繁交替的古气候环境。Cao等(2020)在风城组中发现了大量的可能代表干旱炎热环境的双气囊具肋花粉Striatoabieites和Hamiapollenites,同时仅寻找到代表温暖环境沉积的自生矿物碳镁钠石和碳酸钠钙石,未发现典型的冷水环境沉积矿物水菱镁矿或泡碱,因此认为该组形成于温暖干旱的古气候环境。近期,笔者在玛页1井风城组中新发现了大量孢粉和古鳕类化石,丰富了该组古生物资料,并望能为上述存在争议的风城组地质时代和古气候背景问题提供新的证据。2 地层概况 新疆准噶尔盆地主要由六个一级构造单元组成,包括乌伦古坳陷、陆梁隆起、中央坳陷、西部隆起、东部坳陷和北天山山前冲断带(路玉,2018;王小军等,2021;Wang et al.,2021;Zhi et al.,2021等)。玛湖凹陷位于盆地中央凹陷的西北区,面积约5000 km2,东南与夏盐凸起和达巴松凸起相邻,东北与石英滩凸起和英西凹陷相邻,北以乌夏断裂带为界,西以克百断裂带和中拐凸起东缘为界(何登发等,2018;杨智峰等,2021;Wang et al.,2021)。玛页1井位于玛湖凹陷北部乌夏断裂带内(图1)。玛湖凹陷发育石炭系至第四系,大层系之间多为角度不整合接触(刘得光等,2020;雷海艳等,2022)。本次研究的风城组最初由雍天寿等1983年命名,但在1978年由金玉玕等首次公开发表,命 图 1 玛页 1 井地理位置(底图自路玉,2018 和 Wang et al.,2021,有修改)Fig.1 Location of the Maye-1 borehole in the Mahu sag,northwestern Junggar Basin,Xinjiang,Northwest China(modified from Lu,2018 and Wang et al.,2021)618 古 生 物 学 报 61 卷 名剖面为克拉玛依北侧风城的井下综合剖面,未指定参考剖面(金玉玕等,2000)。风城组为一套泥质、凝灰质白云岩,白云质泥岩夹砂岩、粉砂岩和灰岩薄层,主要分布在乌尔禾夏子街地区(金玉玕等,2000)。风城组的上覆地层为夏子街组,下伏地层为佳木河组,各组之间均为不整合接触(鲁新川等,2012;刘得光等,2020)。玛页1井共计取心444.70 m,主要由风城组组成(康克利等,2020)。玛页1井下部风城组一段(4836.754907.87 m)主要为去云化泥质粉砂岩和流 纹 质 熔 结 角 砾 凝 灰 岩;中 部 风 城 组 二 段(4616.004836.75 m)主要为泥质云岩和云质泥岩互层夹云质粉砂岩;上部风城组三段(4577.67 4616.00 m)主要为泥质白云岩、粉砂质白云岩,夹云质泥岩、粉砂质泥岩(刘占魁,2020)。3 材料和方法 笔者在玛页1井风城组的页岩层中系统采集了50块孢粉化石样品(图2),其中在风城组一段顶部采集了6块孢粉样品,在风城组二段内采集了37块孢粉样品,在风城组三段下部采集了7块孢粉样品。每个样品称取了3050 g碎样,使用标准盐酸氢氟酸盐酸的岩石酸泡方法浸解岩石样品(Wood et al.,和甘油胶固定在生物薄片中。在分析的50个样品中均发现了丰富的孢粉和有机质碎屑。在光学显微镜 图 2 玛页 1 井岩心柱状图和孢粉化石丰度变化图谱 Fig.2 Lithology and percentage changes of palynomorphs of Maye-1 borehole in the Mahu sag,northwestern Junggar Basin,Xinjiang,Northwest China 4期 师天明等:新疆玛湖凹陷孢粉地层学;Shi et al.:Palynostratigraphy of Mahu Sag,Xinjiang 619 1996)。之后,使用孔径180 m和10 m的筛布集中浸解后的有机质碎屑。集中后的有机质碎屑被石蜡下观察,发现孢粉化石普遍经受了较严重的次生蚀变。为了尽可能地获得准确的孢粉统计数据,本文仅对形态保存完好或具有清晰结构的孢粉化石予以鉴定统计(图3)。有机质碎屑以II型干酪根为主,色变指数为2.53.0级,具有很强的生烃潜力。我们使用LEICA DM 2500生物显微镜和Nikon D800E相机对孢粉化石进行了拍照。孢粉固定薄片全部保存在中国科学院南京地质古生物研究所。所有孢粉化石照片号均包含了它们在普通生物显微镜下的坐标位置(England Finder Slide),例如:Z33/1代表孢粉化石的位置在坐标尺第Z行,33列,1数字区。4 风城组孢粉组合特征 玛页1井风城组孢粉组合以双气囊具肋花粉占主导,蕨类孢子和藻类孢子含量极低(表1,图2)。其中,蕨类三缝孢子含量为04.67%,花粉含量为90.33%99.67%,藻类孢子含量为08.67%。三缝孢子类仅个别出现,相对较多的为Leiotriletes or-natus,含量为02.3%;具环孢子Densosporites sp.含量不超过1.7%。在花粉类群中,单气囊或无囊花粉含量为0.3%3%,其中仅Florinites luberae含量稍高,为02%。双气囊无肋花粉含量为3%10.3%,以Alisporites communis为主,含量为05.7%。双气囊花粉含量为88.67%97.33%,其中,双气囊具肋花 粉 含 量 为 81.3%93.3%,以 Protohaploxypinus perfectus占优势(11%27.7%),其次为Lunatisporties tersus(含 量 为 8%22%)和Striatoabieites selliformis(含量为4.7%21.3%)。Hamiapollenites的含量为10%47.9%,平均含量为29.60%,包括H.brachiatus(1.7%14%),H.cf.bullaeformis(5.3%12%),H.fengchengensis(2%11.3%),H.ruiditaeniatus(05.3%)和H.sac-catus(1%5.3%);而双气囊无肋和多气囊无肋花粉 含 量 为 3.33%11%。多 气 囊 花 粉 仅 出 现Triangulisaccites boleensis,含量为01.3%。通过对玛页1井风城组50件样品中孢粉百分含量的聚类分析(CONISS,图2),发现聚类分析的主要分支仅与个别属种类型的突然出现或缺失有关,例如:NIG2-NIG20层段与NIG31-NIG42层段的主要区别是在前者孢粉组合面貌中出现了少量的藻类孢子,除此之外,两者在组合特征方面并没有显示出较大差别。由于玛页1井风城组不同样品中的孢粉化石面貌非常相似,没有出现明显的组分差异,因而可将当前层段产出的孢粉化石划入同一个孢粉组合。由于所有样品中占绝对优势的属种均为双气囊具肋 花 粉,且 Protohaploxypinus perfectus 和 Luna-tisporites tersus 的 含 量 均 较 高,因 而 命 名 为Protohaploxypinus perfectusLunatisporites tersus(PT)孢粉组合。5 对比讨论 5.1 玛页 1 井风城组的地质年代 玛页1井风城组PT孢粉组合以双气囊具肋花粉占主要优势,含量为81.3%93.3%,蕨类孢子含量很低,仅为04.67%。根据欧阳舒等(2017)对新疆北部石炭纪二叠纪裸子植物花粉含量变化的统计分析,当前PT孢粉组合的整体面貌大致与准噶尔盆地南缘奥尔吐组、石人子沟组、塔什库拉组上部、乌拉泊组和井井子沟组的孢粉化石面貌相似,主要体现在出现大量的双气囊具肋花粉(72.3%79.07%)和 含 量 很 低 的 蕨 类 孢 子(0.25%5%)。奥尔吐组产出的Cyclogranisporites aureus Striatoabieites striatusHamiapollenites bullaefor-mis(ASB)孢粉组合(欧阳舒等,2017)中常见的石炭纪 标 志 分 子 Lycospora 和 Noeggerathiopsidozono-triletes并未出现在玛页1井风成组PT孢粉组合中。石人子沟组以砂砾岩为主,其孢粉化石保存普遍较 差,以 Striatoabieites striatusHamiapollenites multistriatus(SM)孢粉组合为代表(欧阳舒等,2017)。SM组合内的双气囊具肋花粉类型和百分含量与PT组合相似,但主要区别在于SM组合中的双囊无肋花粉以Klausipollenites和Pityosporites为代表,而 PT 孢 粉 组 合 中 的 双 囊 无 肋 花 粉 则 以Alisporites communis 为 主,未 发 现 可 靠 的Klausipollenites和Pityosporites。塔什库拉组中、下 620 古 生 物 学 报 61 卷 表1 玛页1井风城组孢粉化石百分含量统计表 Table 1 Percentage chart of palynomorphs of the Fengcheng Formation in Maye-1 borehole from the Mahu sag,northwestern Junggar Basin,Xinjiang,Northwest China 孢粉属种 井深(m)4578.82 4584.09 4598.4 4602.06 4611.67 4619.64 4622.13 4623.16 4631.32 4638.73 4641.36 4662.85 4668.77 4681.47 4699.19 4704.5 4706.79 样品号 NIG334 NIG328 NIG316 NIG310 NIG299 NIG288 NIG285 NIG284 NIG277 NIG270 NIG266 NIG246 NIG237 NIG224 NIG200 NIG191 NIG187 Densosporites sp.0.3 0.3 1 0 0 0 0 0.7 0 0.3 1 0 1 0.7 0.3 0 1 Leiotriletes ornatus 0.7 0.3 0 1 0 0.3 0.3 0.3 1 0.7 0 0 0.7 1 1 0.3 0.7 Retusotriletes levi-densus 0 0.7 1 0.3 0.7 0.3 1.3 0.3 0.7 0.3 1 0 0.7 1 0 0 0 Cordaitina annulata 0.3 0.3 0 0 0.3 1 1.7 1 1.3 0.3 1 0.3 0.3 0.3 0 1.7 0.7 C.subrotata 1 0.3 0.3 0.7 0 0 0 0 0 1.3 0 0 0.7 0.7 1.3 0.3 1 C.uralensis 0 1 1 1.3 1 1 0.7 1.3 0.7 0.3 1.3 1.3 1 1.3 0 0.3 1 Florinites luberae 1 0 1 0.7 1.3 0 0.3 0.3 0.3 0 0 0 0.7 0.3 0.7 0 0 Alisporites communis 0 2.7 2.7 1.3 1.7 0 1.3 2.3 1 3 2.3 1 3 1.7 3.7 2.7 4 A.ovalis 0.7 1 2 1.3 0.3 1.3 1.3 0.3 2 1 0.3 1.3 1 1.3 0.3 1.7 0.7 Cedripites sp.0 0.3 0.3 0.7 0 0 0 0.3 0.3 0 0 0.3 0.7 0.7 0.7 0 0.7 Falcisporites sublevis 1.3 2.3 1.3 1 1.7 1 1.3 1 1.7 0.7 2 1.7 1.7 1 2.3 1.7 2.3 Gardenasporites bilabiatus 2.7 3 2 2 2 1.7 0 1.7 0 0.3 0 1 0 0 1.3 1.7 0 Pinuspollenites solitus 0.7 0 0.3 0 0 0.7 1 0.3 0.3 0.3 0.7 0 0.3 0 0.3 0 0.3 Verrucorpipollis archaicus 0.3 1 0 0 0 1 0.7 1 0.7 0.7 0 0 0 0 1 0 0.3 Hamiapollenites brachiatus 7 7.3 10.7 6.7 11.7 1.7 11 10.3 10.7 8.7 10.3 8.3 9.3 11.7 12.3 5.7 7.7 H.cf.bullaeformis 9.7 9 7 5.3 9 12 7.7 7 10 8.7 7 8.7 9.7 6.3 8 7 8.3 H.fengchengensis 8.3 5.3 7 10 9.3 8 5 8 8.3 5.7 7.3 9 7.3 6.7 5.7 9.7 10.3 H.ruiditaeniatus 0.7 3.3 3.3 2.7 0 1.7 3 3 1.7 3.7 2.7 2.3 1.7 1.3 2 3 2.7 H.saccatus 3.3 1.7 2.3 1.3 2 3.7 2.3 2.7 4 2 2.3 1.7 4 1.7 3.7 2.7 3.7 Lunatisporties tersus 15.7 16.7 14 17.3 15.3 13.7 14 15 13 12.3 13 14.7 16.3 15 11.7 19 15.3 Protohaploxypinus perfectus 19.3 20.7 18.7 23 18.7 21.7 21 20.7 11 18.7 18.3 21 17 17.7 16 20.3 16 P.samoilovichiae 10.3 7.7 8.7 12.7 10.7 11 10.7 6.3 8 12.7 10.3 8 6.7 10.3 10.7 7.3 9 Scutasporites xin-jiangensis 0.3 0 0 0 0.7 0.7 0 1 0 0 0.3 0.3 0 0 0.7 0 0.3 Striatoabieites selliformis 12.3 13 12 8.7 11.7 15.3 10.7 11.7 14.7 15.7 15.3 14 11 14.7 8.7 13.7 8.7 Vittatina specialis 1.7 1 2.7 0.7 0.7 1 2.3 2.3 1.3 2 1.3 3 3 3 1.7 1.3 1.3 Vittatina sp.1 0.3 0 0 0 0.3 0 0.3 0 0.7 0 0.3 0 0.7 0.3 0 0.3 Triangulisaccites boleensis 0 0.7 0 1.3 0 0 0.3 0 1 0 1.3 0 0 1 0.3 0 0.7 Leiosphaeridia sp.1 0 0.7 0 0.3 0 1.3 0.7 4 0 0.3 1.7 1 0 2.3 0 0.7 Rugaletes vietus 0.3 0 0 0 1 1 0.7 0 2.3 0 0.3 0 1.3 0 3 0 2.3 4期 师天明等:新疆玛湖凹陷孢粉地层学;Shi et al.:Palynostratigraphy of Mahu Sag,Xinjiang 621 续表(continued)孢粉属种 井深(m)4711.91 4714.55 4717.24 4725.92 4728.04 4728.59 4740.24 4746.33 4752.32 4755.39 4759.11 4768.06 4770.51 4777.25 4783.39 4783.62 4788.94 样品号 NIG181 NIG177 NIG174 NIG167 NIG164 NIG163 NIG149 NIG142 NIG131 NIG129 NIG122 NIG109 NIG105 NIG99 NIGS42 NIG93 NIG88 Densosporites sp.1 1 0 0.7 0.7 0.3 0.3 0.3 1 0 0.7 0 0.7 0 1 0 0.7 Leiotriletes ornatus 0.3 0.7 0 0.3 0 0 0.7 1 0.7 0.3 1 0 0 0.7 0.3 0.7 0.7 Retusotriletes levi-densus 0.7 0.7 0.3 0 0.7 1 0.7 0.3 0.3 1 0.3 1.3 0.7 0.3 0.3 0.3 0 Cordaitina annulata 1 0.7 1 0.3 0 1 0 0.7 0.3 1.3 0 0 0 0.7 1 0 0 C.subrotata 0 0.7 1.3 1.7 0 0.7 0.3 0 1.7 0 1 0 0.3 0 0 1 0 C.uralensis 0 0 0 0.3 1.7 1 0 0.7 0 0.7 0.3 1.3 1 0 0.3 0 0 Florinites luberae 0 1 0.3 0.7 0 0 1 1.3 0.3 0.3 1 0.7 1 2 0.3 0.7 0.3 Alisporites communis 4 4 2.3 1.7 4 3 2.3 2.7 2 1 4 2.7 2 3 1.3 2.7 1.3 A.ovalis 1.3 0 0 1.3 0.3 0.3 1.3 1.7 1 0.7 0.3 1 1.3 1.3 1 1.3 1 Cedripites sp.0 0 0.7 0.3 0 0 0 0 0.7 0 0.3 0 0 0 0.3 0.3 0 Falcisporites sublevis 1.3 1.7 1.7 2.3 1 2 1.3 1 2.3 1 2.3 1.3 1.3 2.3 0.7 1 0 Gardenasporites bilabiatus 2.3 3 0 0.7 0 2 1.3 0 0 0.7 0 0 0 1.7 0 0.7 1 Pinuspollenites solitus 0 0.3 0 0.3 0.7 0.7 0.7 0.7 0.3 0.3 0.3 0 0.7 0 0.3 0 0 Verrucorpipollis archaicus 0 1 0 0 0.3 1 0 0 0 1 0 0 0.3 0.3 0.7 0 0 Hamiapollenites brachiatus 8.3 7 13.7 8.7 10.3 6.7 9 7.3 13 12.3 9 11.3 7 8.7 7 5.3 8.3 H.cf.bullaeformis 6.7 11 7.3 10 7 9.3 10.3 7.7 6.3 6.3 8 7.3 6.7 7.3 5.3 10 8.3 H.fengchengensis 6.7 9 6 8 4.7 7.3 5.7 6.7 6.3 8.7 6 8.3 7.7 6.3 9.7 5.7 10.7 H.ruiditaeniatus 1.3 1 2 1.3 3.3 3 3.7 2.7 2 4 1.7 3.7 2.3 3 2.7 3 1.7 H.saccatus